Aug 25, 2025  
2025-2026 Franklin & Marshall College Catalog 
  
2025-2026 Franklin & Marshall College Catalog

Animal Behavior Major


Behavior is manifest in the function of neurons, the cells that comprise the nervous system. The networks of a few to many million neurons that underlie the simple and complex behaviors exhibited by humans and animals are shaped by biological, environmental, ecological, evolutionary, social and psychological influences. To develop an understanding of the complex interactions among these factors that generate normal and abnormal behavioral states, critical thinking, reading and writing skills across disciplinary boundaries are required. The Biological Foundations of Behavior Program is offered jointly by the departments of Biology and Psychology. It presents students the opportunity to complete an interdisciplinary major with a focus on either animal behavior or neuroscience.

Neuroscience is an integrative discipline that utilizes knowledge and tools from biology, chemistry, physics, mathematics and psychology to study the nervous system across several levels of analysis from molecules to the behavior of individual organisms. Despite the amazing advances that have been made in neuroscience to date, the human brain is a frontier that we’ve only begun to chart. Understanding how it works, how to protect it from disease and how to fix it when it becomes damaged or diseased is one of humankind’s greatest challenges.

Animal Behavior-Behavior is a fundamental property of all living things. Indeed, whether animals survive and reproduce often depends on how they behave. Studying individual variation in behavior can reveal the role of natural selection in shaping behavior. Comparative research with many species provides animal models for studying development, sensation, perception, life history evolution, reproductive behavior, learning and cognition as well as providing a broader context for better understanding the influences affecting human behavior and the mind. In addition, studying how individuals behave in response to varying environmental conditions can help predict effects of climate change and the fate of populations. Conservation efforts and resource management depend upon ecological and evolutionary studies of the relationship between animal behavior and the environment.

The Neuroscience and the Animal Behavior majors begin with core courses in biology, chemistry, physics and/or mathematics, that create a solid foundation upon which to begin the research-intensive coursework that follows. Following cornerstone courses at the introductory level in neuroscience and biopsychology, Neuroscience students choose elective courses in neuroscience and related areas. After foundational, research-intensive training in animal behavior, Animal Behavior students select from a series of core and elective courses in animal behavior. The Neuroscience and the Animal Behavior majors each culminate with capstone research experiences, typically through independent study, that may be defended for honors in the major during the senior year.

Biological Foundations of Behavior Program Faculty


Professor Ryan T. Lacy, Chair
 

Members of the Biological Foundations of Behavior Committee

Daniel R. Ardia
Professor of Biology

Meredith J. Bashaw
Professor of Psychology

Robert N. Jinks
Professor of Biology

Clara S. Moore
Professor of Biology

Joseph T. Thompson
Professor of Biology

Ryan T. Lacy
Associate Professor of Psychology

Aaron F. Howard
Senior Teaching Professor of Biology

Additional faculty not on the program committee also contribute to this program.

Major Requirements:


A major in Animal Behavior requires the completion of 15 courses:

Research Methods and Statistics (one course)


Fundamentals of Behavior (three courses)


Area Studies Electives (Required of all students. Four courses with no more than two courses chosen from any one area.)


Area 1: Mechanisms of Behavior


Courses that emphasize the neural, endocrine and physiological basis of behavior and cognition.

Note:

Topics courses in neuroscience or biochemistry may serve as Area 1 courses upon approval of the BFB Chair.

Area 2: Organismal and Comparative Approaches


Courses with an emphasis on functional organization and integration within individual organisms.

Note:

Topics courses in biology or psychology may serve as Area 2 courses upon approval of the BFB Chair.

Area 3: Ecological and Population Perspectives


Courses with an emphasis on ecological factors primarily at the population level.

Note:

Topics courses in ecology, environmental studies or ecological psychology may serve as Area 3 courses upon approval of the BFB Chair.

Note:

In Area 4, students may, with permission of the BFB Chair, elect to take a course above the introductory level in a cognate area (e.g., Anthropology, Environmental Studies, Philosophy or Computer Science).

Advanced Research (Required of all students. Take one of the following.)


Note:

An area studies course may not be double-counted as an advanced research course and vice versa.

Note:


To be considered for honors in BFB, graduating seniors, in addition to meeting the College’s general requirements for honors, must possess a cumulative GPA in the major of 3.33 or greater and complete no less than two semesters of independent research in neuroscience or animal behavior. Normally, prospective honors students will enroll in two semesters of BFB 490 .

The writing requirement in the Biological Foundations of Behavior major is met by completion of the normal courses required to complete the major.

The indication as to when a course will be offered is based on the best projection of the BFB Program Committee and the departments of Biology and Psychology and is subject to change

Off-Campus Study:


Majors in the Biological Foundations of Behavior Program have studied abroad in the following programs in recent years: School for Field Studies (Australia, Costa Rica, Kenya, Tanzania); Danish International Study (DIS), Copenhagen; Institute for Study Abroad at Butler University programs (New Zealand, United Kingdom); IES’s Galápagos Islands Direct Enrollment program; Budapest Semester in Cognitive Science; University of Queensland & Macquarie University, Australia. See the International Programs section of the Catalog for further information.